Arthropod P450 Enchiridion

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====== Arthropod P450 sequences ======

For each species, a complete CYPome is provided, obtained by manual curation from genomic and transcriptomic sources. Unless noted otherwise, the CYPomes match a precise genome assembly as stated. Both the sequences and the number of genes are specific for that assembly. [[cnv|Copy number variation]] between strains or populations may result in different CYPome sizes and compositions.

For some species (as indicated below), only partial CYPomes are presented, either from fragmented, low quality genomes, or from transcriptome shotgun assemblies (TSA). The [[https://1kite.org/index.html|1KITE project]] produced a rich and diverse collection of TSAs from insects. 

A few non-arthropod CYPomes (human, yeast, Arabidopsis) are provided for reference [[OTHERcypomes|here]].

All sequences presented here are collected in a file that is searchable by [[https://arthropodp450.eu/sequenceserver/|BLAST]] (see sidebar menu).

The arthropod species are presented according to their phylogenetic position in the tree of life. 

An [[sequences-alpha|alphabetical listing]] of the species is also presented. 

===== Panarthropoda =====

**Onychophora**

  * //[[Euperipatoides|Euperipatoides sp.]]// (partial)

===== Chelicerata =====

**Arachnida** (111 k estimated number of species)

  * //[[Turticae|Tetranychus urticae]]//  
  * //[[Panonychus|Panonychus citri]]//
  * //[[aculops|Aculops lycopersici]]//
  * //[[sarcoptes|Sarcoptes scabiei]]//
  * //[[Dermatophagoides|Dermatophagoides pteronyssinus]]//
  * //[[Dgallinae|Dermanyssus gallinae]]//
  * //[[Metaseiulus|Metaseiulus (Galendromus) occidentalis]]//
  * //[[Varroa|Varroa destructor]]//
  * //[[Stratiolaelaps|Stratiolaelaps scimitus]]//
  * //[[Ixodes|Ixodes scapularis]]//
  * //[[Parasteatoda|Parasteatoda tepidariorum]]// 
  * //[[Mesobuthus|Mesobuthus martensii]]// (partial)
  * //[[Centruroides|Centruroides sculpturatus]]//

**Xiphosura** (5 species)


  * //[[Limulus|Limulus polyphemus]]//


===== Myriapoda =====

**Chilopoda** (centipedes)(15,000 est. species)

  * //[[Strigamia|Strigamia maritima]]//

**Diplopoda** (millipedes)(3,100 est. species)

  * //[[Chamberlinius|Chamberlinius hualienensis]]// (partial)


===== "Crustacea" =====

Crustacea and Hexapoda form a monophyletic clade called Pancrustacea. Within this clade, the crustacea are polyphyletic, but treated here as a group for convenience. See Lozano-Fernandez et al. (2019) for recent phylogeny of [[Crustacea|Crustacea]].

  * //[[Heterocypris|Heterocypris incongruens]]// (partial)
  * //[[Tigriopus|Tigriopus japonicus]]//
  * //[[salmonis|Lepeophtheirus salmonis]]//
  * //[[nordmannii|Lepeophtheirus nordmannii]]//
  * //[[Paracyclopina|Paracyclopina nana]]// (partial - TSA)
  * //[[PVannamei|Penaeus vannamei]]//
  * //[[Hyallella|Hyalella azteca]]//
  * //[[Diaphanosoma|Diaphanosoma celebensis]]//
  * //[[Daphnia|Daphnia pulex]]//

===== Hexapoda =====

For a current overall phylogeny of [[Misof|hexapods and insects]], see the landmark paper of Misof et al. (2014).

==== Protura ====

    * //[[Acerentomon|Acerentomon sp.]]// (partial)
 
==== Collembola ====

    * //[[Folsomia|Folsomia candida]]// 
    * //[[Sinella|Sinella curviseta]]//

==== Diplura ====

    * //[[Catajapyx|Catajapyx aquilonaris]]// (partial)

====  Insecta ====
 (1,054 k est. species)

  * //[[Machilis|Machilis hrabei]]// (partial)
  * //[[Meinertellus|Meinertellus cundinamarcensis]]// (partial - TSA)
  * //[[Thermobia|Thermobia domestica]]// (partial)

**Paleoptera**

  * //[[Calopteryx splendens]]//
  * //[[Ischnura elegans]]//


**Polyneoptera**

  * //[[Locusta|Locusta migratoria]]//
  * //[[Zoraptera TSA]]//
  * //[[Plecoptera TSA]]//
  * //[[Dermaptera TSA]]//
  * //[[Mantophasmatodea TSA]]//
  * //[[Grylloblattodea TSA]]//
  * //[[Embioptera TSA]]//
  * //[[Phasmatodea TSA]]//
  * //[[Mantodea TSA]]//

**Thysanoptera**

**Hemiptera** (104 k est. species)

See Johnson et al. (2018) for a phylogeny of [[Hemiptera]]
  * //[[Nezara viridula]]//
  * //[[Cimex lectularius]]//
  * //[[Rhodnius prolixus]]//
  * //[[Nilaparvata lugens]]//
  * //[[Myzus persicae]]//
  * //[[Acyrthosiphon pisum]]//
  * //[[Bemisia tabaci]]//
  * //[[TSA sequences from other Hemiptera]]//

**Psocodea**

  * //[[Pediculus humanus]]//
 
 
**Hymenoptera** (153 k est. species)

See Branstetter et al. (2017) for a phylogeny of [[Branstetter2017|Aculeate wasps and outgroups]].

  * //[[Neodiprion|Neodiprion lecontei]]//
  * //[[Nasonia vitripennis]]//
  * //[[Ceratosolen solmsi]]//
  * //[[Bombus |Bombus terrestris]]//
  * //[[Apis|Apis mellifera]]//
  * //[[Osmia|Osmia bicornis]]//
  * //[[Megachile|Megachile rotundata]]//

**Raphidioptera-Megaloptera-Neuroptera**

  * // [[Raphidioptera TSA]]// 
  * // [[Megaloptera TSA]]// 
  * // [[Neuroptera TSA]]//

**Strepsiptera**

  * // [[Strepsiptera TSA]]//

**Coleoptera** (392 k est. species)

  * //[[Photinus pyralis]]//
  * //[[Nicrophorus vespilloides]]//
  * //[[Coccinella septempunctata]]//
  * //[[Tribolium|Tribolium castaneum]]//
  * //[[Dendroctonus ponderosae]]//
  * //[[TSA sequences from other Coleoptera]]//

**Diptera** (161 k est. species)

See Wiegmann et al. (2011) for a phylogeny of [[Wiegmann2011|Diptera]].

  * //[[Phlebotomus|Phlebotomus papatasi]]//
  * //[[Lutzomyia|Lutzomyia longipalpis]]//
  * //[[Anopheles|Anopheles gambiae]]//
  * //[[BradysiaC|Bradysia coprophila]]//
  * //[[Hermetia|Hermetia illucens]]//
  * //[[Bactrocera|Bactrocera oleae]]//
  * //[[Sechellia|Drosophila sechellia]]//
  * //[[Drosophila|Drosophila melanogaster]]//

**Mecoptera**

  * // [[ Mecoptera TSA]]// 


**Siphonaptera**

  * //[[catflea|Ctenocephalides felis]]//
  * //[[Siphonaptera TSA]]//

**Trichoptera**

  *//[[Trichoptera TSA]]//

**Lepidoptera** (159 k est. species)

see Kawahara et al. 2019 for a phylogeny of [[ditrysia|Ditrysia]] (higher Lepidoptera)

  * //[[Neomicropteryx|Neomicropteryx cornuta]]//
  * //[[Eumeta|Eumeta japonica]]//
  * //[[Plutella|Plutella xylostella]]//
  * //[[Adoxophyes|Adoxophyes honmai]]//
  * //[[Carposina|Carposina sasakii]]//
  * //[[Zygaena|Zygaena filipendulae]]//
  * //[[Papilio|Papilio xuthus]]//
  * //[[Bicyclus|Bicyclus anynana]]//
  * //[[Pectinophora|Pectinophora gossypiella]]//
  * //[[Pierisr|Pieris rapae]]//
  * //[[Ostrinia|Ostrinia furnacalis]]//
  * //[[Galleria|Galleria mellonella]]//
  * //[[Tni|Trichoplusia ni]]//
  * //[[Slittoralis|Spodoptera littoralis]]//
  * //[[Sfrugiperda|Spodoptera frugiperda]]//
  * //[[Sexigua|Spodoptera exigua]]//
  * //[[Harmigera|Helicoverpa armigera]]//
  * //[[Ectropis|Ectropis grisescens]]//
  * //[[Manduca|Manduca sexta]]//
  * //[[Bombyx|Bombyx mori]]//

**Additional P450 sequences**

[[otherP450s|Named P450 sequences from other arthropod species]]

===== Unusual P450 sequences =====

In addition to CYPomes, several cases of [[StrangeSeqs|unusual P450 sequences]] are described, including insect P450 sequences lacking the [[structure|conserved Cys]], and odd P450 genes that may result from horizontal gene transfer.
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